Friday, February 19, 2010

Aquatic

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Aquatic is a term often used in biology and ecology. Literally, it means watery, and refers to both fresh and salt-water. So aquatic plants or animals are those that live in water: ponds, rivers, oceans. Aquatic mammals include whales, dolphins, seals, walrus, and also otters, beavers, and other river forms. Aquatic birds are even more numerous: ducks, herons, waders, kingfishers, penguins.


Marine biology

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World Marine Environment.

Marine biology is the scientific study of living organisms in the ocean or other marine or brackish bodies of water. Given that in biology many phyla, families and genera have some species that live in the sea and others that live on land, marine biology classifies species based on the environment rather than on taxonomy. Marine biology differs from marine ecology as marine ecology is focused on how organisms interact with each other and environment and biology is the study of the animal itself.

Marine life is a vast resource, providing food, medicine, and raw materials, in addition to helping to support recreation and tourism all over the world. At a fundamental level, marine life helps determine the very nature of our planet. Marine organisms contribute significantly to the oxygen cycle, and are involved in the regulation of the Earth's climate. [1] Shorelines are in part shaped and protected by marine life, and some marine organisms even help create new land.[2]

Marine biology covers a great deal, from the microscopic, including most zooplankton and phytoplankton to the huge cetaceans (whales) which reach up to a reported 48 meters (125 feet) in length.

The habitats studied by marine biology include everything from the tiny layers of surface water in which organisms and abiotic items may be trapped in surface tension between the ocean and atmosphere, to the depths of the abyssal trenches, sometimes 10,000 meters or more beneath the surface of the ocean. It studies habitats such as coral reefs, kelp forests, tidepools, muddy, sandy and rocky bottoms, and the open ocean (pelagic) zone, where solid objects are rare and the surface of the water is the only visible boundary.

A large amount of all life on Earth exists in the oceans. Exactly how large the proportion is unknown, since many ocean species are still to be discovered. While the oceans comprise about 71% of the Earth's surface, due to their depth they encompass about 300 times the habitable volume of the terrestrial habitats on Earth.

Many species are economically important to humans, including food fish. It is also becoming understood that the well-being of marine organisms and other organisms are linked in very fundamental ways. The human body of knowledge regarding the relationship between life in the sea and important cycles is rapidly growing, with new discoveries being made nearly every day. These cycles include those of matter (such as the carbon cycle) and of air (such as Earth's respiration, and movement of energy through ecosystems including the ocean). Large areas beneath the ocean surface still remain effectively unexplored.

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[edit] Subfields

The marine ecosystem is large, and thus there are many subfields of marine biology. Most involve studying specializations of particular animal groups. (i.e. phycology, invertebrate zoology and ichthyology).

Other subfields study the physical effects of continual immersion in sea water and the ocean in general, adaptation to a salty environment, and the effects of changing various oceanic properties on marine life. A subfield of marine biology studies the relationships between oceans and ocean life, and global warming and environmental issues (such as carbon dioxide displacement).

Recent marine biotechnology has focused largely on marine biomolecules, especially proteins, that may have uses in medicine or engineering. Marine environments are the home to many exotic biological materials that may inspire biomimetic materials.

[edit] Related fields

Marine biology is a branch of oceanography and is closely linked to biology. It also encompasses many ideas from ecology. Fisheries science and marine conservation can be considered partial offshoots of marine biology as well as environmental studies.

[edit] Lifeforms

[edit] Microscopic life

Microscopic life undersea is incredibly diverse and still poorly understood. For example, the role of viruses in marine ecosystems is barely being explored even in the beginning of the 21st century.

The role of phytoplankton is better understood due to their critical position as the most numerous primary producers on Earth. Phytoplankton are categorized into cyanobacteria (also called blue-green algae/bacteria), various types of algae (red, green, brown, and yellow-green), diatoms, dinoflagellates, euglenoids, coccolithophorids, cryptomonads, chrysophytes, chlorophytes, prasinophytes, and silicoflagellates.

Zooplankton tend to be somewhat larger, and not all are microscopic. Many Protozoa are zooplankton, including dinoflagellates, zooflagellates, foraminiferans, and radiolarians. Some of these (such as dinoflagellates) are also phytoplankton; the plant/animal distinction often breaks down in very small organisms. Other zooplankton include cnidarians, ctenophores, chaetognaths, molluscs, arthropods, urochordates, and annelids such as polychaetes. Many larger animals begin their life as zooplankton before they become large enough to take their familiar forms. Two examples are fish larvae and sea stars (also called starfish).

[edit] Plants and algae

Plant life is widespread and very diverse under the sea. Microscopic photosynthetic algae contribute a larger proportion of the worlds photosynthetic output than all the terrestrial forests combined. Most of the niche occupied by sub plants on land is actually occupied by macroscopic algae in the ocean, such as Sargassum and kelp, which are commonly known as seaweeds that create kelp forests. The non algae plants that survive in the sea are often found in shallow waters, such as the seagrasses (examples of which are eelgrass, Zostera, and turtle grass, Thalassia). These plants have adapted to the high salinity of the ocean environment. The intertidal zone is also a good place to find plant life in the sea, where mangroves or cordgrass or beach grass might grow. Microscopic algae and plants provide important habitats for life, sometimes acting as hiding and foraging places for larval forms of larger fish and invertebrates.

[edit] Marine invertebrates

As on land, invertebrates make up a huge portion of all life in the sea. Invertebrate sea life includes Cnidaria such as jellyfish and sea anemones; Ctenophora; sea worms including the phyla Platyhelminthes, Nemertea, Annelida, Sipuncula, Echiura, Chaetognatha, and Phoronida; Mollusca including shellfish, squid, octopus; Arthropoda including Chelicerata and Crustacea; Porifera; Bryozoa; Echinodermata including starfish; and Urochordata including sea squirts or tunicates.

[edit] Fish

Fish have evolved very different biological functions from other large organisms. Fish anatomy includes a two-chambered heart, operculum, secretory cells that produce mucous, swim bladder, scales, fins, lips and eyes. Fish breathe by extracting oxygen from water through their gills. Fins propel and stabilize the fish in the water.

Well known fish include: sardines, anchovy, ling cod, clownfish (also known as anemonefish), and bottom fish which include halibut or ling cod. Predators include sharks and barracuda.

[edit] Reptiles

Reptiles which inhabit or frequent the sea include sea turtles, sea snakes, terrapins, the marine iguana, and the saltwater crocodile. Most extant marine reptiles, except for some sea snakes, are oviparous and need to return to land to lay their eggs. Thus most species, excepting sea turtles, spend most of their lives on or near land rather than in the ocean. Despite their marine adaptations, most sea snakes prefer shallow waters not far from land, around islands, especially waters that are somewhat sheltered, as well as near estuaries.[3][4] Some extinct marine reptiles, such as ichthyosaurs, evolved to be viviparous and had no requirement to return to land.

[edit] Seabirds

Seabirds are species of birds adapted to living in the marine environment, examples including albatross, penguins, gannets, and auks. Although they spend most of their lives in the ocean, species such as gulls can often be found thousands of miles inland.

[edit] Marine mammals

There are five main types of marine mammals.

[edit] Oceanic habitats

Ocean habitats

Littoral zone

Intertidal zone

Neritic zone

Continental shelf

Kelp forests

Coral reefs

Ocean banks

Continental margin

Straits

Pelagic zone

Oceanic zone

Seamounts

Hydrothermal vents

Cold seeps

Demersal zone

Benthic zone
Aquatic ecosystems
Aquatic layers
Wild fisheries
Land habitats

[edit] Reefs

Corals and reef fish in Papua New Guinea.

Reefs comprise some of the densest and most diverse habitats in the world. The best-known types of reefs are tropical coral reefs which exist in most tropical waters; however, reefs can also exist in cold water. Reefs are built up by corals and other calcium-depositing animals, usually on top of a rocky outcrop on the ocean floor. Reefs can also grow on other surfaces, which has made it possible to create artificial reefs. Coral reefs also support a huge community of life, including the corals themselves, their symbiotic zooxanthellae, tropical fish and many other organisms.

Much attention in marine biology is focused on coral reefs and the El Niño weather phenomenon. In 1998, coral reefs experienced a "once in a thousand years" bleaching event, in which vast expanses of reefs across the Earth died because sea surface temperatures rose well above normal. Some reefs are recovering, but scientists say that 58% of the world's coral reefs are now endangered and predict that global warming could exacerbate this trend.

[edit] Deep sea and trenches

The deepest recorded oceanic trenches measure to date is the Mariana Trench, near the Philippines, in the Pacific Ocean at 10924 m (35838 ft). At such depths, water pressure is extreme and there is no sunlight, but some life still exists. Small flounder (family Soleidae) fish and shrimp were seen by the American crew of the bathyscaphe Trieste when it dove to the bottom in 1960.

Other notable oceanic trenches include Monterey Canyon, in the eastern Pacific, the Tonga Trench in the southwest at 10,882 m (35,702 ft), the Philippine Trench, the Puerto Rico Trench at 8605 m (28232 ft), the Romanche Trench at 7760 m (24450 ft), Fram Basin in the Arctic Ocean at 4665 m (15305 ft), the Java Trench at 7450 m (24442 ft), and the South Sandwich Trench at 7235 m (23737 ft).

In general, the deep sea is considered to start at the aphotic zone, the point where sunlight loses its power of transference through the water. Many life forms that live at these depths have the ability to create their own light.

Much life centers on seamounts that rise from the depths, where fish and other sea life congregate to spawn and feed. Hydrothermal vents along the mid-ocean ridge spreading centers act as oases, as do their opposites, cold seeps. Such places support unique biomes and many new microbes and other lifeforms have been discovered at these locations.

[edit] Open ocean

The open ocean is relatively unproductive because of a lack of nutrients, yet because it is so vast, in total it produces the most primary productivity. Much of the aphotic zone's energy is supplied by the open ocean in the form of detritus. The open ocean consists mostly of jellyfish and its predators such as the mola mola.

[edit] Intertidal and shore

Tide pools with sea stars and sea anemone in Santa Cruz,California

Intertidal zones, those areas close to shore, are constantly being exposed and covered by the ocean's tides. A huge array of life lives within this zone.

Shore habitats span from the upper intertidal zones to the area where land vegetation takes prominence. It can be underwater anywhere from daily to very infrequently. Many species here are scavengers, living off of sea life that is washed up on the shore. Many land animals also make much use of the shore and intertidal habitats. A subgroup of organisms in this habitat bores and grinds exposed rock through the process of bioerosion.

[edit] Distribution factors

An active research topic in marine biology is to discover and map the life cycles of various species and where they spend their time. Marine biologists study how the ocean currents, tides and many other oceanic factors affect ocean lifeforms, including their growth, distribution and well-being. This has only recently become technically feasible with advances in GPS and newer underwater visual devices.

Most ocean life breeds in specific places, nests or not in others, spends time as juveniles in still others, and in maturity in yet others. Scientists know little about where many species spend different parts of their life cycles. For example, it is still largely unknown where sea turtles and some sharks travel. Tracking devices do not work for some life forms, and the ocean is not friendly to technology. This is important to scientists and fishermen because they are discovering that by restricting commercial fishing in one small area they can have a large impact in maintaining a healthy fish population in a much larger area far away.

[edit] See also


Aquatic ecosystem

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An estuary mouth and coastal waters, part of an aquatic ecosystem.

An aquatic ecosystem is an ecosystem located in a body of water. Communities of organisms that are dependent on each other and on their environment live in aquatic ecosystems. The two main types of aquatic ecosystems are marine ecosystems and freshwater ecosystems.[1]

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[edit] Types

[edit] Marine

Marine ecosystems cover approximately 71% of the Earth's surface and contain approximately 97% of the planet's water. They generate 32% of the world's net primary production.[1] They are distinguished from freshwater ecosystems by the presence of dissolved compounds, especially salts, in the water. Approximately 85% of the dissolved materials in seawater are sodium and chlorine. Seawater has an average salinity of 35 parts per thousand (ppt) of water. Actual salinity varies among different marine ecosystems.[2]

Marine ecosystems can be divided into the following zones: oceanic (the relatively shallow part of the ocean that lies over the continental shelf); profundal (bottom or deep water); benthic (bottom substrates); intertidal (the area between high and low tides); estuaries; salt marshes; coral reefs; and hydrothermal vents (where chemosynthetic sulfur bacteria form the food base).[1]

Classes of organisms found in marine ecosystems include brown algae, dinoflagellates, corals, cephalopods, echinoderms, and sharks. Fish caught in marine ecosystems are the biggest source of commercial foods obtained from wild populations.[1]

Environmental problems concerning marine ecosystems include unsustainable exploitation of marine resources (for example overfishing of certain species), marine pollution, climate change, and building on coastal areas.[1]

[edit] Freshwater

Freshwater ecosystems cover 0.8% of the Earth's surface and contain 0.009% of its total water. They generate nearly 3% of its net primary production.[1] Freshwater ecosystems contain 41% of the world's known fish species.[3]

There are three basic types of freshwater ecosystems:

Lake ecosystems can be divided into zones: pelagic (open offshore waters); profundal; littoral (nearshore shallow waters); and riparian (the area of land bordering a body of water). Two important subclasses of lakes are ponds, which typically are small lakes that intergrade with wetlands, and water reservoirs. Many lakes, or bays within them, gradually become enriched by nutrients and fill in with organic sediments, a process called eutrophication. Eutrophication is accelerated by human activity within the water catchment area of the lake.[1]

Freshwater ecosystem.

The major zones in river ecosystems are determined by the river bed's gradient or by the velocity of the current. Faster moving turbulent water typically contains greater concentrations of dissolved oxygen, which supports greater biodiversity than the slow moving water of pools. These distinctions forms the basis for the division of rivers into upland and lowland rivers. The food base of streams within riparian forests is mostly derived from the trees, but wider streams and those that lack a canopy derive the majority of their food base from algae. Anadromous fish are also an important source of nutrients. Environmental threats to rivers include loss of water, dams, chemical pollution and introduced species.[1]

Wetlands are dominated by vascular plants that have adapted to saturated soil. Wetlands are the most productive natural ecosystems because of the proximity of water and soil. Due to their productivity, wetlands are often converted into dry land with dykes and drains and used for agricultural purposes. Their closeness to lakes and rivers means that they are often developed for human settlement.[1]

[edit] Ponds

These are a specific type of freshwater ecosystems that are largely based on the autotroph algae which provide the base trophic level for all life in the area. The largest predator in a pond ecosystem will normally be a fish and in-between range smaller insects and microorganisms. It may have a scale of organisms from small bacteria to big creatures like water snakes, beetles, water bugs, frogs, tadpoles, and turtles. This is important for the environment.

[edit] Functions

Aquatic ecosystems perform many important environmental functions. For example, they recycle nutrients, purify water, attenuate floods, recharge ground water and provide habitats for wildlife.[5] Aquatic ecosystems are also used for human recreation, and are very important to the tourism industry, especially in coastal regions.[3]

The health of an aquatic ecosystem is degraded when the ecosystem's ability to absorb a stress has been exceeded. A stress on an aquatic ecosystem can be a result of physical, chemical or biological aterations of the environment. Physical alterations include changes in water temperature, water flow and light availability. Chemical alterations include changes in the loading rates of biostimulatory nutrients, oxygen consuming materials, and toxins. Biological alterations include the introduction of exotic species. Human populations can impose excessive stresses on aquatic ecosystems.[5]

[edit] Abiotic characteristics

An ecosystem is composed of biotic communities and abiotic environmental factors, which form a self-regulating and self-sustaining unit. Abiotic environmental factors of aquatic ecosystems include temperature, salinity, and flow.[5]

The amount of dissolved oxygen in a water body is frequently the key substance in determining the extent and kinds of organic life in the water body. Fish need dissolved oxygen to survive. Conversely, oxygen is fatal to many kinds of anaerobic bacteria.[6]

The salinity of the water body is also a determining factor in the kinds of species found in the water body. Organisms in marine ecosystems tolerate salinity, while many freshwater organisms are intolerant of salt. Freshwater used for irrigation purposes often absorb levels of salt that are harmful to freshwater organisms.[6] Though some salt can be good for organisms.

[edit] Biotic characteristics

The organisms (also called biota) found in aquatic ecosystems are either autotrophic or heterotrophic.[6]

[edit] Autotrophic organisms

Autotrophic organisms are producers that generate organic compounds from inorganic material. Algae use solar energy to generate biomass from carbon dioxide and are the most important autotrophic organisms in aquatic environments.[6] Chemosynthetic bacteria are found in benthic marine ecosystems. These organisms are able to feed on hydrogen sulfide in water that comes from volcanic vents. Great concentrations of animals that feed on this bacteria are found around volcanic vents. For example, there are giant tube worms (Riftia pachyptila) 1.5m in length and clams (Calyptogena magnifica) 30cm long.[7]

[edit] Heterotrophic organisms

Heterotrophic organisms consume autotrophic organisms and use the organic compounds in their bodies as energy sources and as raw materials to create their own biomass.[6] Euryhaline organisms are salt tolerant and can survive in marine ecosystems, while stenohaline or salt intolerant species can only live in freshwater environments.[2]

[edit] See also


Aquatic animal

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Longfin Sculpin (Jordania zonope)

An aquatic animal is an animal, either vertebrate or invertebrate, which lives in water for most or all of its life. It may breathe air or extract its oxygen from that dissolved in water through specialised organs called gills, or directly through its skin. Natural environments and the animals that live in them can be categorized as aquatic (water) or terrestrial (land). Animals that move readily from water to land and vice versa are often referred to as amphibious.

The term aquatic can in theory be applied to animals that live in either freshwater or saltwater. However, the adjective marine is most commonly used for animals that live in saltwater, i.e. in oceans, seas, etc.

Aquatic animals (especially freshwater animals) are often of special concern to conservationists because of the fragility of their environments. Aquatic animals are subject to pressure from overfishing, destructive fishing, marine pollution and climate change.

[edit] Air breathing aquatic animals

In addition to water breathing animals,e.g., fishes, mollusks etc., the term "aquatic animal" can be applied to air-breathing aquatic or sea mammals such as those in the order Cetacea (whales), which cannot survive on land, as well as four-footed mammals like the river otter (Lutra canadensis) and beavers (family Castoridae).

Aquatic animals include for example the seabirds, such as gulls (family Laridae), pelicans (family Pelecanidae), and albatrosses (family Diomedeidae), and most of the Anseriformes (ducks, swans and geese).

Amphibious and amphibiotic animals, like frogs (the order Anura), while they do require water, are separated into their own environmental classification. The majority of amphibians (class Amphibia) have an aquatic larval stage, like a tadpole, but then live as terrestrial adults, and may return to the water to mate.

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